| 1970-76 | University of Munich; Study of Biology (Diploma) |
| 1976-81 | University of Munich; Faculty of Biochemistry; PhD thesis in the laboratory of Prof. G. Hartmann |
| 1981-83 | Max-Planck-Institute of Biochemistry, Martinsried; Postdoc in the laboratory of Prof. G. Vogel |
| 1983-91 | University of Munich, Faculty of Medicine; Research assistant in the laboratory of Prof. Walter Neupert; group leader at the Munich Gene Centre |
| since 1991 | University of Freiburg, Faculty of Medicine; Institute for Biochemistry and Molecular Biology; Professor (C3) |
